Citation: | MO Jinyou, WANG Kebai, CHEN Shuqing, WANG Peiye, XU Xing. A new titanosaurian sauropod from the Late Cretaceous strata of Shandong Province[J]. Geological Bulletin of China, 2017, 36(9): 1501-1505. |
Specimen ZJZ-57, a nearly complete sauropod humerus excavated from the Upper Cretaceous Wangshi Group of Zhucheng area, Shandong Province, is described in this paper. Some derived features, such as a lateral bulge at the proximal onethird position of the lateral margin and a deltopectoral crest markedly expanded distally, indicate that ZJZ-57 represents a member of Titanosauria. However, it differs from all other titanosaurian sauropods in the following combined features:the proximal portion is strongly expanded, reaching 55% of the total length of the bone, and the humerus is a robust bone, with a robustness index (RI) of 0.39. Based on this specimen, the authors erect a new sauropod taxon:Zhuchengtitan zangjiazhuangensis gen. et sp. nov. This new taxon is possibly more closely related to Opisthocoelicaudia than to other known titanosaurian sauropods from Asia. This is the first ti-tanosaurian sauropod reported from the Upper Cretaceous strata of Shandong Province.
[1] | 胡承志.山东诸城巨型鸭嘴龙化石[J].地质学报, 1973, (2):179-206. |
[2] | 胡承志, 程政武, 庞其清, 等.巨型山东龙[M].北京:地质出版社, 2001:1-139. |
[3] | 赵喜进, 李敦景, 韩岗, 等.山东的巨大诸城龙[J].地球学报, 2007, 28(2):111-122. |
[4] | 赵喜进, 王克柏, 李敦景.巨大华夏龙[J].地质通报, 2011, 30(11):1671-1688. doi: 10.3969/j.issn.1671-2552.2011.11.003 |
[5] | Xu X, Wang K B, Zhao X J, et al. First ceratopsid dinosaur from China and its biogeographical implications[J]. Chinese Science Bulletin, 2010, 55(16):1631-1635. doi: 10.1007/s11434-009-3614-5 |
[6] | Xu X, Wang K B, Zhao X J, et al. A new leptoceratopsid(Ornithischia:Ceratopsia) from the Upper Cretaceous of Shandong, China and its implications for neoceratopsian evolution[J]. Plos One, 2010, 5(11):e13825. doi: 10.1371/journal.pone.0013825 |
[7] | Hone D W E, Wang K B, Sullivan C, et al. A new, large tyrannosaurine theropod from the Upper Cretaceous of China[J]. Cretaceous Research, 2011, 32(4):495-503. doi: 10.1016/j.cretres.2011.03.005 |
[8] | He Y M, Makovicky P J, Wang K B, et al. A new leptoceratopsid (Ornithischia, Ceratopsia) with a unique ischium from the Upper Cretaceous of Shandong Province, China[J]. Plos One, 2015, 10(12):e0144148. doi: 10.1371/journal.pone.0144148 |
[9] | Li L, Tong H Y, Wang K B, et al. Lindholmemydid turtles(Cryptodira:Testudinoidea) from the Late Cretaceous of Shandong Province, China[J]. Annales de Paléontologie, 2013, 99(3):243-259. doi: 10.1016/j.annpal.2013.07.003 |
[10] | Wiman C. Die Kreide-Dinosaurier aus Shantung[M]. Paleontologica Sinica, Series C, 1929, 6(1):1-67. |
[11] | 杨钟健. 山东莱阳恐龙化石[C]//中国古生物志(142册), 新丙种(16). 北京: 科学出版社, 1958: 1-138. |
[12] | 董枝明.山东莱阳王氏组中一肿头龙[J].古脊椎动物与古人类, 1978, 16(4):225-228. |
[13] | Buffetaut E, Tong H Y. The Late Cretaceous dinosaurs of Shandong, China:old finds and new interpretations[C]//Sun A L, Wang Y Q. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota. Beijing:China Ocean Press, 1995:139-142. |
[14] | 杨钟健.山东莱阳蛋化石[J].古生物学报, 1954, 2:371-388. |
[15] | 赵资奎, 蒋元凯.山东莱阳恐龙蛋化石的显微结构研究[J].中国科学, 1974, (1):63-77. |
[16] | 刘金远, 赵资奎.山东莱阳晚白垩世恐龙蛋化石一新类型[J].古脊椎动物学报, 2004, 42:166-170. doi: 10.3969/j.issn.1000-3118.2004.02.008 |
[17] | 柳永清, 旷红伟, 彭楠, 等.鲁东诸城地区晚白垩世恐龙集群埋藏地沉积相与埋藏学初步研究[J].地质论评, 2010, 56(4):457-468. |
[18] | Gilmore C W. Reptilian fauna of the North Horn Formation of central Utah[J]. United States Geological Survey Professional Paper, 1946, 210-C:29-51. |
[19] | Lawson D A. Paleoecology of the Tornillo Formation, Big Bend National Park, Brewster County, Texas[D]. Unpublished M.A. thesis, University of Texas at Austin, 1972:1-182. |
[20] | Lehman T M, Coulson A B. A juvenile specimen of the sauropod dinosaur Alamosaurus sanjuanensis from the Upper Cretaceous of Big Bend National Park, Texas[J]. Journal of Paleontology, 2002, 76:156-172. doi: 10.1666/0022-3360(2002)076<0156:AJSOTS>2.0.CO;2 |
[21] | Huene F Von. Los Saurisquiosy Ornithisquios de Cretaceo Argentino[M]. Anales del Museo de La Plata(series 2), 1929, 3:1-196. |
[22] | Powell J E. Revision de los titanosauridos de America del Sur[D]. Unpublished Ph.D. dissertation, Universidad Nacional de Tucuman, Argentina, 1986:1-493. |
[23] | Powell J E. Osteología de Saltasaurus loricatus(Sauropoda-Titanosauridae) del Cretácico superior des noroeste Argentino[C]//Sanz J L, Buscalioni A D. Los Dinosaurios y su Entornoe Biotico. Actas del Segundo Curso de Paleontología en Cuenca, Instituto "Juan de Valdes".Cuenca, Spain, 1992:165-230. |
[24] | Borsuk-Bialynicka M. A new camarasaurid sauropod, Opisthocoelicaudia skarzynski gen. n. sp. n. from the Upper Cretaceous of Mongolia[J]. Palaeontologia Polonica, 1977, 37:1-64. |
[25] | 莫进尤, 黄超林, 赵仲如, 等.中国广西晚白垩世一新的巨龙类恐龙[J].古脊椎动物学报, 2008, 46(2):147-156. |
[26] | Pang Q Q, Cheng Z W. A new family of sauropod dinosaur from the Upper Cretaceous of Tianzhen, Shanxi Province, China[J]. Acta Geologica Sinica, 2000, 74(2):117-125. |
[27] | D' Emic M D, Mannion P D, Upchurch P, et al. Osteology of Huabeisaurus allocotus (Sauropoda:Titanosauriformes) from the Upper Cretaceous of China[J]. Plos One, 2013, 8(8):e69375. doi: 10.1371/journal.pone.0069375 |
[28] | Wilson J A. Sauropod dinosaur phylogeny:critique and cladistic analysis[J]. Zoological Journal of the Linnean Society, 2002, 136:217-276. |
[29] | D' Emic M D. The early evolution of titanosauriform sauropod dinosaurs[J]. Zoological Journal of the Linnean Society, 2012, 166:624-671. doi: 10.1111/zoj.2012.166.issue-3 |
[30] | You H L, Ji Q, Matthew C L, et al. A titanosaurian sauropod dinosaur with opisthocoelous caudal vertebrae from the early Late Cretaceous of Liaoning Province, China[J]. Acta Geologica Sinica, 2004, 78(4):907-911. |
[31] | Martin V, Buffetaut E, Suteethorn V. A new genus of sauropod di-nosaur from the Sao Khua Formation(Late Jurassic or Early Cretaceous) of Northeastern Thailand[J]. Comptes Rendus de l' Académie des Sciences de Paris, 1994, 319(2):1085-1092. |
[32] | Martin V, Suteethorn V, Buffetaut E. Description of the type and referred material of Phuwiangosaurus sirindhornae Martin, Buffetaut and Suteethorn, 1994, a sauropod from the Lower Cretaceous of Thailand[J]. Oryctos, 1999, 2:39-91. |
Holotype (ZJZ-57) of Zhuchengtitan zangjiazhuangensis
Humeral morphology in several Late Cretaceous titanosaurian sauropods